Fig. 1

Territorial separation of two individuals (or individual packs) on a one-dimensional landscape in avoidance of scent marks deposited by one another in the environment. Den sites are located at the two ends of the landscape. The yellow and green lines represent the individual UDs; their sum is shown in black. The respective individual scent-mark densities are displayed by the red and blue lines. The process of spatial segregation during the initial 10,000 time steps post-release is captured by the transient UD dynamics; mutually deterrent boundary markings continue to accumulate long after space-use convergence