Fig. 1
From: Identifying signals of memory from observations of animal movements

Spatial memory can be quantified using an animal’s occurrence distribution (OD) measured over some prior period (colored areas: 5-day periods at time 6 and time 8 = orange vs green) which captures an animal’s movement path and its uncertainty. A time-varying covariate can be constructed by updating the OD at regular time intervals. This updating step ensures that distant experiences are eventually forgotten and no longer play a role in driving animal movement. Users must choose an appropriate time window in the past for calculating the memory covariate and how often to update it